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Like previous Halo games, Halo: Reach has difficulty-multipliers in the appearance of human skulls. In addition, all but one of the skulls featured in Halo 3 are included in Reach. Unlike Halo 3, you do not need to find and unlock these skulls in order to use them; they are all already unlocked when you start playing Reach.
In Firefight, you may have up to 3 different custom skulls, which you can change things such as enemy firing style, luck, and also player traits.