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One player's shields recharge to overshield level and can only melee while the second player has no shields but infinite ammo.
Masterblaster is a skull in Halo 2: Anniversary.
In co-op, one of the players has an Overshield but can only Melee, whilst the other player doesn't have shields at all, but has infinite ammo. These roles switch after a certain number of kills made by the players.